Table 1 Five existing TFBS-based RS measures

Garten et al.¹

$-\mathsf{\text{log}}\left({\displaystyle \sum _{x\ge k}^{\mathsf{\text{min}}\left(m,n\right)}}\frac{\left(\begin{array}{c}\hfill m\hfill \\ \hfill x\hfill \end{array}\right)\left(\begin{array}{c}\hfill N-m\hfill \\ \hfill n-x\hfill \end{array}\right)}{\left(\begin{array}{c}\hfill N\hfill \\ \hfill n\hfill \end{array}\right)}\right)$ Eq. (1)

Veerla and Höglund

$\frac{\left|T{F}_a\cap T{F}_b\right|}{\left|T{F}_a\cup T{F}_b\right|}$ Eq. (2)

Shalgi et al.

$\frac{\left|T{F}_a\cap T{F}_b\right|}{\mathsf{\text{min}}\left(\left|T{F}_a\right|,\left|T{F}_b\right|\right)}$ Eq. (3)

Park et al.²

$\begin{array}{cc}S\hfill & ={\displaystyle \sum _{j=1}^2}{\gamma }_j{\displaystyle \sum _i}{\left(f_i^j\right)}^{-1/2}\hfill \\ \left[\left(\frac{2}{N_{1i}^j+N_{2i}^j}+\alpha \right)C_i^j-\beta \left(N_{1i}^j+N_{2i}^j\right)I_{\left\{C_i^j=0\right\}}\right]\hfill \end{array}$ Eq. (4)

van Helden²

$M^{ab}=S^{ab}-\alpha D^{ab}+\beta$ Eq. (5)